Neurotransmitters Definition and List

Neurotransmitters are chemicals that cross synapses to transmit impulses from a neuron to another neuron, glandular cell, or muscle cell. In other words, neurotransmitters are used to send signals from one part of the body to another. Over 100 neurotransmitters are known. Many are simply constructed from amino acids. Others are more complex molecules. Neurotransmitters perform many vital functions in the body. For example, they regulate heartbeat, tell the lungs when to breathe, determine the set point for weight, stimulate thirst, affect mood, and control digestion. The synaptic cleft was discovered by Spanish pathologist  Santiago Ramà ³n y Cajal in the early 20th century. In 1921, German pharmacologist Otto Loewi verified that communication between neurons was the result of released chemicals. Loewi discovered the first known neurotransmitter, acetylcholine. How Neurotransmitters Work The axon terminal of a synapse stores neurotransmitters in vesicles. When stimulated by an action potential, synaptic vesicles of a synapse release neurotransmitters, which cross the small distance (synaptic cleft) between an axon terminal and a dendrite via diffusion. When the neurotransmitter binds a receptor at the dendrite, the signal is communicated. The neurotransmitter remains in the synaptic cleft for a short time. Then it is either returned to the presynaptic neuron through the process of reuptake, metabolized by enzymes, or bound to the receptor. When a neurotransmitter binds to a postsynaptic neuron, it can either excite it or inhibit it. Neurons are often connected to other neurons, so at any given time a neuron  may be subject to multiple neurotransmitters. If the stimulus for excitation is greater than the inhibitory effect, the neuron will fire and create an action potential that releases neurotransmitters to another neuron. Thus, a signal is conducted from one cell to the next. Types of Neurotransmitters One method of classifying neurotransmitters is based on their chemical composition. Categories include: Amino acids: ÃŽ ³-aminobutyric acid (GABA), aspartate, glutamate, glycine, D-serineGases: carbon monoxide (CO), hydrogen sulfide (H2S), nitric oxide (NO)Monoamines: dopamine, epinephrine, histamine, norepinephrine, serotoninPeptides:  ÃŽ ²-endorphin, amphetamines, somatostatin, enkephalinPurines: adenosine, adenosine triphosphate (ATP)Trace amines: octopamine, phenethylamine, trypramineOther molecules: acetylcholine, anandamideSingle ions: zinc The other major method of categorizing neurotransmitters is according to whether they are excitatory or inhibitory. However, whether a neurotransmitter is excitatory or inhibitory depends on its receptor. For example, acetylcholine is inhibitory to the heart (slows heart rate), yet excitatory to skeletal muscle (causes it to contract). Important Neurotransmitters Glutamate is the most abundant neurotransmitter in humans, used by about half of the neurons in the human brain. It is the primary excitatory transmitter in the central nervous system. One of its functions is to help form memories. Interestingly, glutamate is toxic to neurons. Brain damage or a stroke can lead to an excess of glutamate, killing neurons.GABA is the primary inhibitory transmitter in the vertebrate brain. It helps to control anxiety. GABA deficiency may result in seizures.Glycine is the main inhibitory neurotransmitter in the vertebrate spinal cord.Acetylcholine stimulates muscles, functions in the autonomic nervous system and sensory neurons, and is associated with REM sleep. Many poisons act by blocking acetylcholine receptors. Examples include botulin, curare, and hemlock. Alzheimers disease is associated with a significant drop in acetylcholine levels.Norepinephrine (noradrenaline) increases heart rate and blood pressure. It is part of the bodys fight or flight syst em. Norepinephrine is also needed to form memories. Stress depletes stores of this neurotransmitter.Dopamine is an inhibitory transmitter associated with the reward center of the brain. Low dopamine levels are associated with social anxiety and Parkinsons disease, while excess dopamine is related to schizophrenia.Serotonin is an inhibitory neurotransmitter involved in mood, emotion, and perception. Low serotonin levels can lead to depression, suicidal tendencies, anger management issues, difficulty sleeping, migraines, and an increased craving for carbohydrates. The body can synthesize serotonin from the amino acid tryptophan, which is found in foods such as warm milk and turkey.Endorphins are a class of molecules similar to opioids (e.g., morphine, heroin) in terms of structure and function. The word endorphin is short for endogenous morphine. Endorphins are inhibitory transmitters associated with pleasure and pain relief. In other animals, these chemicals slow metabolism and permi t hibernation.

The Brave New World By Aldous Huxley - 952 Words

Truth vs. Happiness â€Å"If one’s different, one’s bound to be lonely†(Huxley 90), stated the savage John in Brave New World, by Aldous Huxley . In this novel, the government banned all kinds of human truth like friendship, love, books, and any emotions. The citizen cannot know the truth, because if they found out the truth, they will not trust science and the government. The citizens are fed with lies in order for the government to keep them happy and distracted from the truth, but it’s not real happiness. The Resident world controller Mustapha Mond says happiness is more important, when actually the truth is; the citizens are brainwashed with lies and told to take soma. In this essay we’ll be discussing about the isolation and compatibility of truth vs. happiness. To have a utopian society in Brave New World the government achieves a state of stability, and gets rid of freedom. The government places the citizens in a caste system before they are even born and in the test tube alcohol is injected. Babies aren’t naturally born from a mother’s womb; and the words mother and father aren’t used in this world. One egg is divided eight then into ninety-six to even the society out. To not face racism, skin tone difference and stereotypes, but in Utopia that’s eliminated from society. Everyone belongs to everyone else, meaning there’s no relationship or marriage at all. A lot of sex and drugs happen in the book and the people all believe in the same god â€Å"Henry Ford†. The citizensShow MoreRelatedA Brave New World by Aldous Huxley668 Words   |  3 PagesIn Brave New World, there are similarities that have a deeper meaning that we can understand. There are personal effects in Aldous H uxley life that contribute to what he has written in the book. Aldous Huxley throughout his life have seen, done, and events have happened to him, just like all of us, but he has expressed it in his book. So when Aldous wrote the he had so many ideas. I have read the book; it’s notRead MoreBrave New World by Aldous Huxley811 Words   |  3 Pages Brave New World is based around characters who gave up the right of freedom for happiness; characters who ignored the truth so that they could live in a utopian civilization. The deceiving happiness was a constant reminder throughout the book. Almost every character in Brave New World did whatever they could to avoid facing the truth about their own situations. In this society, happiness is not compatible with the truth because the World State believes that happiness was at the expense of theRead MoreBrave New World By Aldous Huxley1525 Words   |  7 PagesA Brave New Feminist The novel Brave New World written by Aldous Huxley in 1932 is known for its social satire, utopian values, and unusual standpoints on stereotypical gender roles. In this time where futuristic technology has completely taken over, and men and women are given the same opportunities for everything, â€Å"the genders appear equal within the social order; both men and women work at the same jobs, have equal choice in sexual partners, and participate in the same leisure pursuits† (MarchRead MoreA Brave New World by Aldous Huxley664 Words   |  3 Pagesfor the fact being in the future and in the past time has changed and many differences were made. In his Dystopian Society Huxley portrays masses of niches where the government produces clones for specific reasons. Huxley decides throughout Brave New World that cloning humans is unethical. He then becomes in contact with the society’s most powerful Alphas and Betas clones. Huxley suggest in BNW that lower class groups in clo ning humans to act like servants to terrorize them into working hard conditionsRead MoreA Brave New World by Aldous Huxley1189 Words   |  5 Pages In the world of sex, drugs, and baby cloning you are going to be in many situations where you feel like the world we live in should be different. In the story Brave New World, they had sex with multiple partners along with a very bad use of drugs. It is weird that Aldous Huxley wrote this book in 1931 about the world he was living in during that time and how it is similar to the world we live in today. Nowadays, drugs are still being used and people are still engaging in sexual encounters withRead MoreBrave New World By Aldous Huxley968 Words   |  4 PagesAldous Huxley’s utopia in Brave New World foreshadowed and illuminated the complications within modern day society. Upon its release, the narrative became widely banned all over the United States due to the unorthodox thoughts and actions of multiple characters in it. Early readers, as well as modern day audiences, feared and rejected the ideals that Huxley incorporated into his perfect society; however, our society today is heading towards the dark paths the older generations desired to avoid. Read MoreA Brave New World by Aldous Huxley895 Words   |  4 Pagesthe novel Brave New World, Aldous Huxley tells of a society where everyone is the same but, compared to t oday’s society, everything is different. Huxley tells of a world where everything that happens or takes place is because of one’s own desire and nothing more. The hero in the novel, a â€Å"savage† named John, is Huxley’s main focal point. It is through his eyes and mind that the reader sees what’s going on. Now when I read this novel, I began to think, â€Å"Could this perfect, conformed world actuallyRead MoreA Brave New World by Aldous Huxley1684 Words   |  7 Pagesimperfect world and is usually only a hopeful dream. These types of worlds can greatly be described in detail through the world of science fiction. Aldous Huxley was an English writer who lived during a time when war and chaos were engulfing the world. His works reflect his view and thoughts on a dystopia, which is a false utopia, and describes what could occur in possible governments of the world. The ability to understand and dive into the thoughts of the author is what make s world literatureRead MoreA Brave New World by Aldous Huxley614 Words   |  2 Pagesthem truly happy. What if someone were to tell you that what you thought was true happiness was all an illusion. In a Brave New World by Aldous Huxley people in the world state are conditioned and drugged up by soma to not experience true happiness. In a world that is perfect, human beings do not have to depend on drugs to keep our world in balance. In a Brave New World by Aldous Huxley there is always a perfect drug called soma that keeps everyone happy, which they have based their society on. ThisRead MoreA Brave New World by Aldous Huxley948 Words   |  4 PagesWelcome to a world were â€Å"Brave† is not just a word; It has a true meaning. This is a story were everything as you know it, doesn’t seem to be right and will completely change your way of thinking. When this story was written, life was very harsh for many people†¦.Mostly for the author who wrote â€Å"Brave New World† During this time (1930s) they didn’t have much sexual content Living The Future Of The Past In The Present†¦.. In the air; But Aldous made a future full of sex for them and we are the

Wireless Sensor Networks Free Essays

1. Introduction The increasing interest in wireless sensor networks can be promptly understood simply by thinking about what they essentially are: a large number of small sensing self-powered nodes which gather information or detect special events and communicate in a wireless fashion, with the end goal of handing their processed data to a base station. Sensing, processing and communication are three key elements whose combination in one tiny device gives rise to a vast number of applications [A1], [A2]. We will write a custom essay sample on Wireless Sensor Networks or any similar topic only for you Order Now Sensor networks provide endless opportunities, but at the same time pose formidable challenges, uch as the fact that energy is a scarce and usually non-renewable resource. However, recent advances in low power VLSI, embedded computing, communication hardware, and in general, the convergence of computing and communications, are making this emerging technology a reality [A3]. Likewise, advances in nanotechnology and Micro Electro-Mechanical Systems (MEMS) are pushing toward networks of tiny distributed sensors and actuators. 2. Applications of Sensor Networks Possible applications of sensor networks are of interest to the most diverse fields. Environmental monitoring, warfare, child education, surveillance, micro-surgery, and griculture are only a few examples [A4]. Through joint efforts of the University of California at Berkeley and the College of the Atlantic, environmental monitoring is carried out off the coast of Maine on Great Duck Island by means of a network of Berkeley motes equipped with various sensors [B6]. The nodes send their data to a base station which makes them available on the Internet. Since habitat monitoring is rather sensitive to human presence, the deployment of a sensor network provides a noninvasive approach and a remarkable degree of granularity in data acquisition [B7]. The same idea lies behind the Pods project at the University of Hawaii at Manoa [B8], where environmental data (air temperature, light, wind, relative humidity and rainfall) are gathered by a network of weather sensors embedded in the communication units deployed in the South-West Rift Zone in Volcanoes National Park on the Big Island of Hawaii. A major concern of the researchers was in this case camouflaging the sensors to make them invisible to curious tourists. In Princeton’s Zebranet Project [B9], a dynamic sensor network has been created by attaching special collars equipped with a low-power GPS system to the necks of zebras to onitor their moves and their behavior. Since the network is designed to operate in an infrastructure-free environment, peer-to-peer swaps of information are used to produce redundant databases so that researchers only have to encounter a few zebras in order to collect the data. Sensor networks can also be used to monitor and study natural phenomena which intrinsically discourag e human presence, such as hurricanes and forest fires. Joint efforts between Harvard University, the University of New Hampshire, and the University of North Carolina have recently led to the deployment of a wireless sensor etwork to monitor eruptions at Volcan Tungurahua, an active volcano in central Ecuador. A network of Berkeley motes monitored infrasonic signals during eruptions, and data were transmitted over a 9 km wireless link to a base station at the volcano observatory [B10]. Intel’s Wireless Vineyard [B11] is an example of using ubiquitous computing for agricultural monitoring. In this application, the network is expected not only to collect and interpret data, but also to use such data to make decisions aimed at detecting the presence of parasites and enabling the use of the appropriate kind of insecticide. Data collection relies on data mules, small devices carried by people (or dogs) that communicate with the nodes and collect data. In this project, the attention is shifted from reliable information collection to active decisionmaking based on acquired data. Just as they can be used to monitor nature, sensor networks can likewise be used to monitor human behavior. In the Smart Kindergarten project at UCLA [B12], wirelessly-networked, sensor-enhanced toys and other classroom objects supervise the learning process of children and allow unobtrusive monitoring by the teacher. Medical research and healthcare can greatly benefit rom sensor networks: vital sign monitoring and accident recognition are the most natural applications. An important issue is the care of the elderly, especially if they are affected by cognitive decline: a network of sensors and actuators could monitor them and even assist them in their daily routine. Smart appliances could help them organize their lives by remindin g them of their meals and medications. Sensors can be used to capture vital signs from patients in real-time and relay the data to handheld computers carried by medical personnel, and wearable sensor nodes can store patient data such as identification, history, and treatments. With these ideas in mind, Harvard University is cooperating with the School of Medicine at Boston University to develop CodeBlue, an infrastructure designed to support wireless medical sensors, PDAs, PCs, and other devices that may be used to monitor and treat patients in various medical scenarios [B13]. On the hardware side, the research team has Martin Haenggi is with the Department of Electrical Engineering, University of Notre Dame, Notre Dame, IN 46556; Fax +1 574 631 4393; mhaenggi@@nd. edu. Daniele Puccinelli is also with the Department of Electrical Engineering, University of Notre Dame, Notre Dame, IN 46556. reated Vital Dust, a set of devices based on the MICA21 sensor node platform (one of the most popular members of the Berkeley motes family), which collect heart rate, oxygen saturation, and EKG data and relay them over a medium-range (100 m) wireless network to a PDA [B14]. Interactions between sensor networks and humans are already judged controversial. The US has recen tly approved the use of a radio-frequency implantable device (VeriChip) on humans, whose intended application is accessing the medical records of a patient in an emergency. Potential future repercussions of this decision have been discussed in the media. An interesting application to civil engineering is the idea of Smart Buildings: wireless sensor and actuator networks integrated within buildings could allow distributed monitoring and control, improving living conditions and reducing the energy consumption, for instance by controlling temperature and air flow. Military applications are plentiful. An intriguing example is DARPA’s self-healing minefield [B15], a selforganizing sensor network where peer-to-peer communication between anti-tank mines is used to respond to attacks and redistribute the mines in order to heal breaches, complicating the progress of enemy troops. Urban warfare is another application that distributed sensing lends itself to. An ensemble of nodes could be deployed in a urban landscape to detect chemical attacks, or track enemy movements. PinPtr is an ad hoc acoustic sensor network for sniper localization developed at Vanderbilt University [B16]. The network detects the muzzle blast and the acoustic shock wave that originate from the sound of gunfire. The arrival times of the acoustic events at different sensor nodes are used to estimate the position of the sniper and send it to the base station with a special data aggregation and routing service. Going back to peaceful applications, efforts are underway at Carnegie Mellon University and Intel for the design of IrisNet (Internet-scale Resource-Intensive Sensor Network Services) [B17], an architecture for a worldwide sensor web based on common computing hardware such as Internet-connected PCs and low-cost sensing hardware such as webcams. The network interface of a PC indeed senses the virtual environment of a LAN or the Internet rather than a physical environment; with an architecture based on the concept of a distributed database [B18], this hardware can be orchestrated into a global sensor system hat responds to queries from users. 3. Characteristic Features of Sensor Networks In ad hoc networks, wireless nodes self-organize into an infrastructureless network with a dynamic topology. Sensor networks (such as the one in Figure 1) share these traits, but also have several distinguishing features. The number of nodes in a typical sensor network is much higher than in a typical ad hoc network, and dense deployments are often desired to ensure coverage and connectivity; for these reasons, sensor network hardware must be cheap. Nodes typically have stringent energy limitations, which make them more failure-prone. They are enerally assumed to be stationary, but their relatively frequent breakdowns and the volatile nature of the wireless channel nonetheless result in a variable network topology. Ideally, sensor network hardware should be power-efficient, small, inexpensive, and reliable in order to maximize network lifetime, add flexibility, facilitate data collection and minimize the need for maintenance. Lifetime Lifetime is extremely critical for most applications, and its primary limiting factor is the energy consumption of the nodes, which need to be self-powering. Although it is often assumed that the transmit power associated with acket transmission accounts for the lion’s share of power consumption, sensing, signal processing and even hardware o peration in standby mode consume a consistent amount of power as well [C19], [C20]. In some applications, extra power is needed for macro-scale actuation. Many researchers suggest that energy consumption could be reduced by considering the existing interdependencies between individual layers in the network protocol stack. Routing and channel access protocols, for instance, could greatly benefit from an information exchange with the physical layer. At the physical layer, benefits can be obtained with ower radio duty cycles and dynamic modulation scaling (varying the constellation size to minimize energy expenditure THIRD QUARTER 2005 IEEE CIRCUITS AND SYSTEMS MAGAZINE 21 External Infrastructure Gateway Base Station Sensing Nodes Figure 1. A generic sensor network with a two-tiered archi1 tecture. See Section 5 for a hardware overview. [D35]). Using low-power modi for the processor or disabling the radio is generally advantageous, even though periodically turning a subsystem on and of f may be more costly than always keeping it on. Techniques aimed at reducing the idle mode leakage current in CMOS-based rocessors are also noteworthy [D36]. Medium Access Control (MAC) solutions have a direct impact on energy consumption, as some of the primary causes of energy waste are found at the MAC layer: collisions, control packet overhead and idle listening. Powersaving forward error control techniques are not easy to implement due to the high amount of computing power that they require and the fact that long packets are normally not practical. Energy-efficient routing should avoid the loss of a node due to battery depletion. Many proposed protocols tend to minimize energy consumption on forwarding aths, but if some nodes happen to be located on most forwarding paths (e. g. , close to the base station), their lifetime will be reduced. Flexibility Sensor networks should be scalable, and they should be able to dynamically adapt to changes in node density and topology, like in the case of the self-healing minefields. In surveillance applications, most nodes may remain quiescent as long as nothing interesting happens. However, they must be able to respond to special events that the network intends to study with some degree of granularity. In a self-healing minefield, a number of sensing mines ay sleep as long as none of their peers explodes, but need to quickly become operational in the case of an enemy attack. Response time is also very critical in control applications (sensor/actuator networks) in which the network is to provide a delay-guaranteed service. Untethered systems need to self-configure and adapt to different conditions. Sensor networks should also be robust to changes in their topology, for instance due to the failure of individual nodes. In particular, connectivity and coverage should always be guaranteed. Connectivity is achieved if the base station can be reached from any node. Coverage can be seen as a measure of quality of service in a sensor network [C23], as it defines how well a particular area can be observed by a network and characterizes the probability of detection of geographically constrained phenomena or events. Complete coverage is particularly important for surveillance applications. Maintenance The only desired form of maintenance in a sensor network is the complete or partial update of the program code in the sensor nodes over the wireless channel. All sensor nodes should be updated, and the restrictions on the size of the new code should be the same as in the case of wired programming. Packet loss must be accounted for and should not impede correct reprogramming. The portion of code always running in the node to guarantee reprogramming support should have a small footprint, and updating procedures should only cause a brief interruption of the normal operation of the node [C24]. The functioning of the network as a whole should not be endangered by unavoidable failures of single nodes, which may occur for a number of reasons, from battery depletion to unpredictable external events, and may either be independent or spatially correlated [C25]. Faulttolerance is particularly crucial as ongoing maintenance s rarely an option in sensor network applications. Self-configuring nodes are necessary to allow the deployment process to run smoothly without human interaction, which should in principle be limited to placing nodes into a given geographical area. It is not desirable to have humans configure nodes for habitat monitoring and destructively interfere with wildlife in the process, or configure nodes for urban warfare monitoring in a hostile environment. The nodes should be able to assess the quality of the network deployment and indicate any problems that may arise, as well as adjust to hanging environmental conditions by automatic reconfiguration. Location awareness is important for selfconfiguration and has definite advantages in terms of routing [C26] and security. Time synchronization [C27] is advantageous in promoting cooperation among nodes, such as data fusion, channel access, coordination of sleep modi, or security-related interaction. Data Collection Data collection is related to network connectivity and coverage. An interesting solution is the use of ubiquitous mobile agents that randomly move around to gather data bridging sensor nodes and access points, whimsically named data MULEs (Mobile Ubiquitous LAN Extensions) in [C28]. The predictable mobility of the data sink can be used to save power [C29], as nodes can learn its schedule. A similar concept has been implemented in Intel’s Wireless Vineyard. It is often the case that all data are relayed to a base station, but this form of centralized data collection may shorten network lifetime. Relaying data to a data sink causes non-uniform power consumption patterns that may overburden forwarding nodes [C21]. This is particularly harsh on nodes providing end links to base stations, which may end up relaying traffic coming from all ther nodes, thus forming a critical bottleneck for network throughput [A4], [C22], as shown in Figure 2. An interesting technique is clustering [C30]: nodes team up to form clusters and transmit their information to their cluster heads, which fuse the data and forward it to a 22 IEEE CIRCUITS AND SYSTEMS MAGAZINE THIRD QUARTER 2005 sink. Fewer packets are transmitted, and a un iform energy consumption pattern may be achieved by periodic re-clustering. Data redundancy is minimized, as the aggregation process fuses strongly correlated measurements. Many applications require that queries be sent to sensing nodes. This is true, for example, whenever the goal is gathering data regarding a particular area where various sensors have been deployed. This is the rationale behind looking at a sensor network as a database [C31]. A sensor network should be able to protect itself and its data from external attacks, but the severe limitations of lower-end sensor node hardware make security a true challenge. Typical encryption schemes, for instance, require large amounts of memory that are unavailable in sensor nodes. Data confidentiality should be preserved by encrypting data with a secret key shared with the intended receiver. Data integrity should be ensured to revent unauthorized data alteration. An authenticated broadcast must allow the verification of the legitimacy of data and their sender. In a number of commercial applications, a serious disservice to the user of a sensor network is compromising data availability (denial of service), which can be achieved by sleep-deprivation torture [C33]: batte ries may be drained by continuous service requests or demands for legitimate but intensive tasks [C34], preventing the node from entering sleep modi. 4. Hardware Design Issues In a generic sensor node (Figure 3), we can identify a power module, a communication block, a processing unit ith internal and/or external memory, and a module for sensing and actuation. Power Using stored energy or harvesting energy from the outside world are the two options for the power module. Energy storage may be achieved with the use of batteries or alternative devices such as fuel cells or miniaturized heat engines, whereas energy-scavenging opportunities [D37] are provided by solar power, vibrations, acoustic noise, and piezoelectric effects [D38]. The vast majority of the existing commercial and research platforms relies on batteries, which dominate the node size. Primary (nonrechargeable) batteries are often chosen, predominantly AA, AAA and coin-type. Alkaline batteries offer a high energy density at a cheap price, offset by a non-flat discharge, a large physical size with respect to a typical sensor node, and a shelf life of only 5 years. Voltage regulation could in principle be employed, but its high inefficiency and large quiescent current consumption call for the use of components that can deal with large variations in the supply voltage [A5]. Lithium cells are very compact and boast a flat discharge curve. Secondary (rechargeable) batteries are typically not desirable, as they offer a lower energy density and a higher cost, not to mention the fact that in most pplications recharging is simply not practical. Fuel cells [D39] are rechargeable electrochemical energy- conversion devices where electricity and heat are produced as long as hydrogen is supplied to react with oxygen. Pollution is minimal, as water is the main byproduct of the reaction. The potential of fuel cells for energy storage and power del ivery is much higher than the one of traditional battery technologies, but the fact that they require hydrogen complicates their application. Using renewable energy and scavenging techniques is an interesting alternative. Communication Most sensor networks use radio communication, even if lternative solutions are offered by laser and infrared. Nearly all radio-based platforms use COTS (Commercial Off-The-Shelf) components. Popular choices include the TR1000 from RFM (used in the MICA motes) and the CC1000 from Chipcon (chosen for the MICA2 platform). More recent solutions use industry standards like IEEE 802. 15. 4 (MICAz and Telos motes with CC2420 from Chipcon) or pseudo-standards like Bluetooth. Typically, the transmit power ranges between ? 25 dBm and 10 dBm, while the receiver sensitivity can be as good as ? 110 dBm. THIRD QUARTER 2005 IEEE CIRCUITS AND SYSTEMS MAGAZINE 23 Base Station Critical Nodes Figure 2. A uniform energy consumption pattern should avoid the depletion of the resources of nodes located in the vicinities of the base station. Communication Hardware Power Sensors (? Actuators) ADC Memory Processor Figure 3. Anatomy of a generic sensor node. Spread spectrum techniques increase the channel reliability and the noise tolerance by spreading the signal over a wide range of frequencies. Frequency hopping (FH) is a spread spectrum technique used by Bluetooth: the carrier frequency changes 1600 times per second on the basis of a pseudo-random algorithm. However, channel synchronization, hopping sequence search, and the high data rate ncrease power consumption; this is one of the strongest caveats when using Bluetooth in sensor network nodes. In Direct Sequence Spread Spectrum (DSSS), communication is carried out on a single carrier frequency. The signal is multiplied by a higher rate pseudo-random sequence and thus spread over a wide frequency range (typical DSSS radios have sprea ding factors between 15 and 100). Ultra Wide Band (UWB) is of great interest for sensor networks since it meets some of their main requirements. UWB is a particular carrier-free spread spectrum technique where the RF signal is spread over a spectrum as large as several GHz. This implies that UWB signals look like noise to conventional radios. Such signals are produced using baseband pulses (for instance, Gaussian monopulses) whose length ranges from 100 ps to 1 ns, and baseband transmission is generally carried out by means of pulse position modulation (PPM). Modulation and demodulation are indeed extremely cheap. UWB provides built-in ranging capabilities (a wideband signal allows a good time resolution and therefore a good location accuracy) [D40], allows a very low power consumption, and performs well in the presence of multipath fading. Radios with relatively low bit-rates (up to 100 kbps) re advantageous in terms of power consumption. In most sensor networks, high data rates are not needed, even though they allow shorter transmission times thus permitting lower duty cycles and alleviating channel access contention. It is also desirable for a radio to quickly switch from a sleep mode to an operational mode. Optical transceivers such as lasers offer a strong power advantage, mainly due to their high directionality and the fact that only baseband processing is required. Also, security is intrinsically guaranteed (intercepted signals are altered). However, the need for a line of sight and recise localization makes this option impractical for most applications. Processing and Computing Although low-power FPGAs might become a viable option in the near future [D41], microcontrollers (MCUs) are now the primary choice for processing in sensor nodes. The key metric in the selection of an MCU is power consumption. Sleep modi deserve special attention, as in many applications low duty cycles are essential for lifetime extension. Just as in the case of the radio module, a fast wake-up time is important. Most CPUs used in lower-end sensor nodes have clock speeds of a few MHz. The memory requirements depend on the pplication and the network topology: data storage is not critical if data are often relayed to a base station. Berkeley motes, U CLA’s Medusa MK-2 and ETHZ’s BTnodes use low-cost Atmel AVR 8-bit RISC microcontrollers which consume about 1500 pJ/instruction. More sophisticated platforms, such as the Intel iMote and Rockwell WINS nodes, use Intel StrongArm/XScale 32-bit processors. Sensing The high sampling rates of modern digital sensors are usually not needed in sensor networks. The power efficiency of sensors and their turn-on and turn-off time are much more important. Additional issues are the physical ize of the sensing hardware, fabrication, and assembly compatibility with other components of the system. Packaging requirements come into play, for instance, with chemical sensors which require contact with the environment [D42]. Using a microcontroller with an onchip analog comparator is another energy-saving technique which allows the node to avoid sampling values falling outside a certain range [D43]. The ADC which complements analog sensors is particularly critical, as its resolution has a direct impact on energy consumption. Fortunately, typical sensor network applications do not have stringent resolution requirements. Micromachining techniques have allowed the miniaturization of many types of sensors. Performance does decrease with sensor size, but for many sensor network applications size matters much more than accuracy. Standard integrated circuits may also be used as temperature sensors (e. g. , using the temperaturedependence of subthreshold MOSFETs and pn junctions) or light intensity transducers (e. g. , using photodiodes or phototransistors) [D44]. Nanosensors can offer promising solutions for biological and chemical sensors while concurrently meeting the most ambitious miniaturization needs. 5. Existing Hardware Platforms Berkeley motes, made commercially available by Crossbow, are by all means the best known sensor node hardware implementation, used by more than 100 research organizations. They consist of an embedded microcontroller, low-power radio, and a small memory, and they are powered by two AA batteries. MICA and MICA2 are the most successful families of Berkeley motes. The MICA2 platform, whose layout is shown in Figure 4, is equipped with an Atmel ATmega128L and has a CC1000 transceiver. A 51-pin expansion connector is available to interface sensors (commercial sensor boards designed for this specific platform are available). Since the MCU is to handle 24 IEEE CIRCUITS AND SYSTEMS MAGAZINE THIRD QUARTER 2005 medium access and baseband processing, a fine-grained event-driven real-time operating system (TinyOS) has been implemented to specifically address the concurrency and resource management needs of sensor nodes. For applications that require a better form factor, the circular MICA2Dot can be used: it has most of the resources of MICA2, but is only 2. 5 cm in diameter. Berkeley motes up to the MICA2 generation cannot interface with other wireless- enabled devices [E47]. However, the newer generations MICAz and Telos support IEEE 802. 15. , which is part of the 802. 15 Wireless Personal Area Network (WPAN) standard being developed by IEEE. At this point, these devices represent a very good solution for generic sensing nodes, even though their unit cost is still relatively high (about $100–$200). The proliferation of different lowerend hardware platforms within the Berkeley mote family has recently led to the development of a new version of TinyOS which introduces a flexible hardware abstraction architecture to simplify multi-platform support [E48]. Tables 1 and 2 show an overview of the radio transceivers and the microcontrollers most commonly used in xisting hardware platforms; an overview of the key features of the platforms is provided in Table 3. Intel has designed its own iMote [E49] to implement various improvements over available mote designs, such as increased CPU processing power, increased main memory size for on-board computing and improved radio reliability. In the iMote, a powerful ARM7TDMI core is complemented by a large main memory and non-volatile storage area; on the radio side, Bluetooth has been chosen. Various platforms have been developed for the use of Berkeley motes in mobile sensor networks to enable investigations into controlled mobility, which facilitates eployment and network repair and provides possibilities for the implementation of energy-harve sting. UCLA’s RoboMote [E50], Notre Dame’s MicaBot [E51] and UC Berkeley’s CotsBots [E52] are examples of efforts in this direction. UCLA’s Medusa MK-2 sensor nodes [E53], developed for the Smart Kindergarten project, expand Berkeley motes with a second microcontroller. An on-board power management and tracking unit monitors power consumption within the different subsystems and selectively powers down unused parts of the node. UCLA has also developed iBadge [E54], a wearable sensor node with sufficient computational power to process the sensed data. Built around an ATMega128L and a DSP, it features a Localization Unit designed to estimate the position of iBadge in a room based on the presence of special nodes of known location attached to the ceilings. In the context of the EYES project (a joint effort among several European institutions) custom nodes [E55], [C24] have been developed to test and demonstrate energy-efficient networking algorithms. On the software side, a proprietary operating system, PEEROS (Preemptive EYES Real Time Operating System), has been implemented. The Smart-Its project has investigated the possibility of embedding computational power into objects, leading o the creation of three hardware platforms: DIY Smartits, Particle Computers and BTnodes. The DIY Smart-its [E56] have been developed in the UK at Lancaster University; their modular design is based on a core board that provides processing and communication and can be extended with add-on boards. A typical setup of Smart-its consists of one or more sen sing nodes that broadcast their data to a base station which consists of a standard core board connected to the serial port of a PC. Simplicity and extensibility are the key features of this platform, which has been developed for the creation of Smart Objects. An interesting application is the Weight Table: four load cells placed underneath a coffee table form a Wheatstone bridge and are connected to a DIY node that observes load changes, determines event types like placement and removal of objects or a person moving a finger across the surface, and also retrieves the position of an object by correlating the values of the individual load cells after the event type (removed or placed) has been recognized [E57]. Particle Computers have been developed at the University of Karlsruhe, Germany. Similarly to the DIY platform, the Particle Smart-its are based on a core board quipped with a Microchip PIC; they are optimized for energy efficiency, scalable communication and small scale (17 mm ? 30 mm). Particles communicate in an ad hoc fashion: as two Particles come close to one another, THIRD QUARTER 2005 IEEE CIRCUITS AND SYSTEMS MAGAZINE 25 Oscillator 7. 3728-MHz DS2401P Silicon Serial No. Antenna Connector Connector LEDs Battery Connection 32. 768-kHz Oscillator 14. 7456-MHz Oscillator ATMEL ATMega 128L CPU CC1000 Transceiver ATMEL AT45DB041 Data Flash Figure 4. Layout of the MICA2 platform. they are able to talk. Additionally, if Particles come near a gateway device, they can be connected to Internet-enabled evices and access services and information on the Internet as well as provide information [E58]. The BTnode hardware from ETHZ [E47] is based on an Atmel ATmega128L microcontroller and a Bluetooth module. Although advertised as a low-power technology, Bluetooth has a relatively high power consumption, as discussed before. It also has long connection setup times and a lower degree of freedom with respect to possible network topologies. On the other hand, it ensures interoperability between different devices, enables application development through a standardized interface, and offers a significantly higher bandwidth (about 1 Mbps) ompared to many low-power radios (about 50 Kbps). Moreover, Bluetooth support means that COTS hardware can be used to create a gateway between a sensor network and an external network (e. g. , the Internet), as opposed to more costly proprietary solutions [E59]. MIT is working on the ? AMPS (? -Adaptive Multidomain Power-aware Sensors) project, which explores energy-efficiency constraints and key issues such as selfconfiguration, reconfigurability, and flexibility. A first prototype has been designed with COTS components: three stackable boards (processing, radio and power) and an ptional extension module. The energy dissipation of this microsensor node is reduced through a variety of poweraware design techniques [D45] including fine-grain shutdown of inactive components, dynamic voltage and frequency scaling of the processor core, and adjustable radio transmission power based on the required range. Dynamic voltage scaling is a technique used for active power management where the supply voltage and clock frequency of the processor are regulated depending on the computa tional load, which can vary significantly based on the operational mode [D36], [C20]. The main oal of second generation ? AMPS is clearly stated in [D46] as breaking the 100 ? W average power barrier. Another interesting MIT project is the Pushpin computing system [E60], whose goal is the modelling, testing, and deployment of distributed peer-to-peer sensor networks consisting of many identical nodes. The pushpins are 18 mm ? 18 mm modular devices with a power substrate, an infrared communication module, a processing module (Cygnal C8051F016) and an expansion module (e. g. , for sensors); they are powered by direct contact between the power substrate and layered conductive sheets. 26 MCU Max. Freq. [MHz] Memory Data Size [bits] ADC [bits] Architecture AT90LS8535 (Atmel) 4 8 kB Flash, 512B EEPROM, 512B SRAM 8 10 AVR ATMega128L (Atmel) 8 128 kB Flash, 4 kB EEPROM, 4 kB SRAM 8 10 AVR AT91FR4081 (Atmel) 33 136 kB On-Chip SRAM, 8 Mb Flash 32 — Based on ARM core (ARM7TDMI) MSP430F149 (TI) 8 60 kB + 256B Flash, 2 kB RAM 16 12 Von Neumann C8051F016 (Cygnal) 25 2304B RAM, 32 kB Flash 8 10 Harvard 8051 PIC18F6720 (Microchip) 25 128 kB Flash, 3840B SRAM, 1 kB EEPROM 8 10 Harvard PIC18F252 (Microchip) 40 32 K Flash, 1536B RAM, 256B EEPROM 8 10 Harvard StrongARM SA-1110 (Intel) 133 — 32 — ARM v. 4 PXA255 (Intel) 400 32 kB Instruction Cache, 32 kB Data 32 — ARM v. 5TE Cache, 2 kB Mini Data Cache Table 2. Microcontrollers used in sensor node platforms. Radio (Manufacturer) Band [MHz] Max. Data Rate [kbps] Sensit. [dBm] Notes TR1000 (RFM) 916. 5 115. 2 ? 106 OOK/ASK TR1001 (RFM) 868. 35 115. 2 ? 106 OOK/ASK CC1000 (Chipcon) 300–1,000 76. 8 ? 110 FSK, ? 20 to 10 dBm CC2420 (Chipcon) 2,400 250 ? 94 OQPSK, ? 24 to 0 dBm, IEEE 802. 15. 4, DSSS BiM2 (Radiometrix) 433. 92 64 ? 93 9XStream (MaxStream) 902–928 20 ? 114 FHSS Table 1. Radios used in sensor node platforms. IEEE CIRCUITS AND SYSTEMS MAGAZINE THIRD QUARTER 2005 MIT has also built Tribble (Tactile reactive interface built by linked elements), a spherical robot wrapped by a wired skinlike sensor network designed to emulate the functionalities of biological skin [E61]. Tribble’s surface is divided into 32 patches with a Pushpin processing module and an array of sensors and actuators. At Lancaster University, surfaces provide power and network connectivity in the PinPlay project. Network nodes come in different form factors, but all share the PinPlay connector, a custom component that allows physical connection and networking through conductive sheets which re embedded in surfaces such as a wall or a bulletin board [E62]. PinPlay falls in between wired and wireless technologies as it provides network access and power across 2D surfaces. Wall-mounted objects are especially suited to be augmented to become PinPlay objects. In a demonstration, a wall switch was augmented and freely placed anywhere on a wall with a PinPlay surface as wallpap er. For applications which do not call for the minimization of power consumption, high-end nodes are available. Rockwellis WINS nodes and Sensoria’s WINS 3. 0 Wireless Sensing Platform are equipped with more powerful rocessors and radio systems. The embedded PC modules based on widely supported standards PC/104 and PC/104-plus feature Pentium processors; moreover, PC/104 peripherals include digital I/O devices, sensors and actuators, and PC-104 products support almost all PC software. PFU Systems’ Plug-N-Run products, which feature Pentium processors, also belong to this category. They offer the capabilities of PCs and the size of a sensor node, but lack built-in communication hardware. COTS components or lower-end nodes may be used in this sense [C32]. Research is underway toward the creation of sensor nodes that are more capable than the motes, yet maller and more power-efficient than higher-end nodes. Simple yet effective gateway devices are the MIB programming boar ds from Crossbow, which bridge networks of Berkeley motes with a PC (to which they interface using the serial port or Ethernet). In the case of Telos motes, any generic node (i. e. , any Telos mote) can act as a gateway, as it may be connected to the USB port of a PC and bridge it to the network. Of course, more powerful gateway devices are also available. Crossbow’s Stargate is a powerful embedded computing platform (running Linux) with enhanced communication and sensor signal processing capabilities based n Intel PXA255, the same X-Scale processor that forms the core of Sensoria WINS 3. 0 nodes. Stargate has a connector for Berkeley motes, may be bridged to a PC via Ethernet or 802. 11, and includes built-in Bluetooth support. 6. Closing Remarks Sensor networks offer countless challenges, but their versatility and their broad range of applications are eliciting more and more interest from the research community as well as from industry. Sensor networks have the potential of triggering the next revolution in information technology. The challenges in terms of circuits and systems re numerous: the development of low-power communication hardware, low-power microcontrollers, MEMSbased sensors and actuators, efficient AD conversion, and energy-scavenging devices is necessary to enhance the potential and the performance of sensor networks. System integration is another major challenge that sensor networks offer to the circuits and systems research community. We believe that CAS can and should have a significant impact in this emerging, exciting area. 27 Platform CPU Comm. External Memory Power Supply WesC (UCB) AT90LS8535 TR1000 32 kB Flash Lithium Battery MICA (UCB, Xbow) ATMega128L TR1000 512 kB Flash AA MICA2 (UCB, Xbow) ATMega128L CC1000 512 kB Flash AA MICA2Dot (UCB, Xbow) ATMega128L CC1000 512 kB Flash Lithium Battery MICAz (UCB, Xbow) ATMega128L CC2420 512 kB Flash AA Telos (Moteiv) MSP430F149 CC2420 512 kB Flash AA iMote (Intel) ARM7TDMI Core Bluetooth 64 kB SRAM, 512 kB Flash AA Medusa MK-2 (UCLA) ATMega103L TR1000 4 Mb Flash Rechargeable Lithium Ion AT91FR4081 iBadge (UCLA) ATMega128L Bluetooth, TR1000 4 Mb Flash Rechargeable Lithium Ion DIY (Lancaster University) PIC18F252 BiM2 64 Kb FRAM AAA, Lithium, Rechargeable Particle (TH) PIC18F6720 RFM TR1001 32 kB EEPROM AAA or Lithium Coin Battery or Rechargeable BT Nodes (ETHZ) ATMega128L Bluetooth, CC1000 244 kB SRAM AA ZebraNet (Princeton) MSP430F149 9XStream 4 Mb Flash Lithium Ion Pushpin (MIT) C8051F016 Infrared — Power Substrate WINS 3. 0 (Sensoria) PXA255 802. 11b 64 MB SDRAM, 32 MB + 1 GB Flash Batteries Table 3. Hardware features of various platforms. THIRD QUARTER 2005 IEEE CIRCUITS AND SYSTEMS MAGAZINE Acknowledgments The support of NSF (grants ECS 03-29766 and CAREER CNS 04-47869) is gratefully acknowledged. References General References [A1] I. F. Akyildiz, W. Su, Y. Sankarasubramaniam, and E. Cayirci, â€Å"A survey on sensor networks,† in IEEE Communications Magazine, pp. 02–114, Aug. 2002. [A2] L. B. Ruiz, L. H. A. Correia, L. F. M. Vieira, D. F. Macedo, E. F. Nakamura, C. M. S. Figueiredo, M. A. M. Vieira, E. H. B. Maia, D. Camara, A. A. F. Loureiro, J. M. S. Nogueira, D. C. da Silva Jr. , and A. O. Fernandes, â€Å"Architectures for wireless sensor networks (In Portuguese),† in Proceedings of the 22nd Brazilian Symposium on Computer Networks (SBRC’04), Gramado, Brazil, pp. 167–218, May 2004. Tutorial. ISBN: 85-88442-82-5. [A3] C. Y. Chong and S. P. Kumar, â€Å"Sensor networks: Evolution, opportunities, and challenges,† in IEEE Proceedings, pp. 1247–1254, Aug. 003. [A4] M. 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Thiele, â€Å"Prototyping sensor network applications with BTnodes,† in IEEE European Workshop on Wireless Sensor Networks (EWSN’04), Berlin, Germany, Jan. 2004. [E60] J. Lifton, D. Seetharam, M. Broxton, an d J. Paradiso, â€Å"Pushpin computing system overview: A platform for distributed, embedded, ubiquitous sensor networks,† in Proceedings of the Pervasive Computing Conference, Zurich, Switzerland, Aug. 2002. [E61] J. A. Paradiso, J. Lifton, and M. Broxton, â€Å"Sensate media—multimodal electronic skins as dense sensor networks,† BT Technology Journal, vol. 2, pp. 32–44, Oct. 2004. [E62] K. V. Laerhoven, N. Villar, and H. -W. Gellersen, â€Å"PinMix: When Pins Become Interaction Components. . . ,† in Physical Interaction (PI03)— Workshop on Real World User Interfaces†Ã¢â‚¬â€Mobile HCI Conference, Udine, Italy, Sept. 2003. Daniele Puccinelli received a Laurea degree in Electrical Engineering from the University of Pisa, Italy, in 2001. After spending two years in industry, he joined the graduate program in Electrical Engineering at the University of Notre Dame, and received an M. S. Degree in 2005. He is currently working toward his P h. D. degree. His research has focused on cross-layer approaches to wireless sensor network protocol design, with an emphasis on the interaction between the physical and the network layer. Martin Haenggi received the Dipl. Ing. (M. Sc. ) degree in electrical engineering from the Swiss Federal Institute of Technology in Zurich (ETHZ) in 1995. In 1995, he joined the Signal and Information Processing Laboratory at ETHZ as a Teaching and Research Assistant. In 1996 he earned the Dipl. NDS ETH (post-diploma) degree in information technology, and in 1999, he completed his Ph. D. thesis on the analysis, design, and optimization of ellular neural networks. After a postdoctoral year at the Electronics Research Laboratory at the University of California in Berkeley, he joined the Department of Electrical Engineering at the University of Notre Dame as an assistant professor in January 2001. For both his M. Sc. and his Ph. D. theses, he was awarded the ETH medal, and he received an NSF CAREER award in 2005. F or 2005/06, he is a CAS Distinguished Lecturer. His scientific interests include networking and wireless communications, with an emphasis on ad hoc and sensor networks. THIRD QUARTER 2005 IEEE CIRCUITS AND SYSTEMS MAGAZINE 29 How to cite Wireless Sensor Networks, Papers